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After graduating from college, Andrea "Andy" Sachs (Anne Hathaway) lands the dream job that "a million girls would kill for": assistant to Miranda Priestly (Meryl Streep) - the chief editor of Runway, the top-selling fashion magazine in the industry. It is a job set to fast track her career in journalism if she can survive a year working for Miranda. From here, Andy, with no sense of fashion at all, begins a fish-out-of-water drama as she is thrown into a lifestyle full of the fast-paced, three-inch-minimum heel height, diet Coke and coffee substance abuse. Andy works really hard to deal with Miranda's endless unimaginable demands. She even becomes trendy and classy. However, she gradually finds she is working 24/7, and soon her life with boyfriend, Nate (Adrian Grenier), and best friend, Lily (Tracie Thoms), is slipping away from her. Then, she realizes that she is losing what really matters. She does not want to lose herself no matter how many pairs of Monolos and Jimmy Choos she can score along the way.